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Basic
Description of Lipids and Lipoproteins
Cholesterol is a fat-like
substance (lipid) that is present in cell membranes and is a precursor
of bile acids and steroid hormones. Cholesterol travels in the blood in
distinct particles containing both lipid and proteins. These particles
are called lipoproteins. The cholesterol level in the blood is determined
partly by inheritance and partly by acquired factors such as diet, calorie
balance, and level of physical activity.
Lipoproteins are a
class of heterogeneous particles of varying sizes and densities containing
lipid and protein. These lipoproteins include cholesterol esters and free
cholesterol, triglycerides, phospholipids and A, C, and E apolipoproteins.
Total cholesterol comprises all the cholesterol found in various lipoproteins.
Factors that affect blood cholesterol levels include age, sex, body weight,
diet, alcohol and tobacco use, exercise, genetic factors, family history,
medications, menopausal status, the use of hormone replacement therapy,
and chronic disorders such as hypothyroidism, obstructive liver disease,
pancreatic disease (including diabetes), and kidney disease. In many individuals,
an elevated blood cholesterol level constitutes an increased risk of developing
coronary artery disease. Blood levels of total cholesterol and various
fractions of cholesterol, especially low density lipoprotein cholesterol
(LDL-C) and high density lipoprotein cholesterol (HDL- C), are useful
in assessing and monitoring treatment for patients with cardiovascular
and related diseases. Blood levels of the above cholesterol components
including triglyceride have been separated into desirable, borderline
and high risk categories by the National Heart, Lung and Blood Institute
in their report in 1993. These categories form a useful basis for evaluation
and treatment of patients with hyperlipidemia (See NCEP Guidelines). Therapy
to reduce these risk parameters includes diet, exercise and medication,
and fat weight loss, which is particularly powerful when combined with
diet or exercise.
Three major classes
of lipoproteins are found in the blood of a fasting individual: low density
lipoproteins (LDL), high density lipoproteins (HDL), and very low density
lipoproteins (VLDL). The LDL typically contain 50-70 percent of the total
serum cholesterol and both are directly correlated with risk for CHD.
The HDL normally contain 20-30 percent of the total cholesterol, and HDL
levels are inversely correlated with CHD risk. The VLDL contain 10-15
percent of the total serum cholesterol along with most of the triglyceride
in fasting serum; VLDL are precursors of LDL, and some forms of VLDL,
particularly VLDL remnants, appear to be atherogenic.
Lesser amounts of
cholesterol are also carried in two minor lipoprotein classes, intermediate
density lipoproteins (IDL, d 1.006-1.019 kg/L) and Lp(a) (d 1.045-1.080
kg/L).
Chylomicrons (d <1.006
kg/L) appear in the blood transiently after a fat-containing meal and
are normally removed completely by 12 hours. They are rich in triglycerides
and responsible for the postprandial increase in plasma triglycerides
but usually have no significant effect on the total cholesterol concentration.
The major lipoprotein fractions can be readily separated from each other,
in the ultracentrifuge and are most commonly quantified in terms of their
cholesterol .
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